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Biological Sciences

Master: José Quintans, BSLC 104A, 702-7964, qui4@midway.uchicago.edu
Senior Adviser: Manfred D. E. Ruddat, HM 254, 702-8623, mdr4@midway.uchicago.edu
Administrative Assistant: G. Karen Shrode, BSLC 104C, 702-7962, shrode@midway.uchicago.edu
Staff Secretary: Cynthia Piwowar, BSLC 104B, 702-7963, piwo@midway.uchicago.edu
Faculty Advisers: James Hopson, Organismal Biology & Anatomy, A 405B, 702-8099; Laurens Mets, Molecular Genetics & Cell Biology, EBC 210, 702-8917; Michael Wade, Ecology & Evolution, Z 14, 702-6830; Marvin Makinen, Biochemistry & Molecular Biology, CLSC 359,
702-1080; Harry Fozzard, Pharmacological & Physiological Sciences, AMB M603, 702-1481; Dorothy Hanck, Neuroscience Concentration, AMB M604A, 702-1758
Undergraduate Research and Honors: Wolfgang Epstein, CLSC 449,
702-1331; Summer Undergraduate Research: James Miller, CLSC 1021, 702-0981; Minority Undergraduate Research: Terence Martin, CLSC 739, 702-8043

Program of Study

Biology is the study of living things and their adaptations to the pressures of natural selection. The faculty of the College believes that a sound knowledge of biology is essential for understanding many of the most pressing problems of modern life and for intelligent involvement in their eventual solution. The Biological Sciences Collegiate Division, therefore, provides a variety of Common Core courses for all College students, prospective biologists and nonbiologists alike. Although most of the course offerings beyond the introductory year are designed to serve the needs of biology concentrators, many of these courses are well suited to students in other concentrations who wish to study some aspect of modern biology in greater detail. Courses on the ethical and societal implications of the biological sciences, for example, will be of interest to all students.

The Biological Sciences Common Core

The goal of the courses in the Common Core biological sciences curriculum is to give students a scientific understanding of how life functions and evolves at every level. Biology is complex, fascinating, and of increasing importance as the "biological revolution" unfolds. Common Core sequences cover the biology of cells, organisms, and populations. Students learn the key concepts and methods to generate new knowledge in each of these areas, and come to appreciate the interconnectedness of all living things and their evolutionary interactions with the environment.

There are three categories of sequences, which are described in greater detail in the course descriptions of this section. These include (1) an integrated six-quarter natural science sequence, which fulfills both the Common Core biological and physical sciences requirements; (2) biological sciences sequences for students who plan to concentrate outside the biological sciences; and (3) advanced biological science sequences designed both to fulfill the Common Core requirement and to prepare biological sciences concentrators for further work in the biological sciences concentration.

Each Common Core course includes laboratory exercises. In these laboratories, students gain firsthand experience in carrying out experiments or making observations and in analyzing and interpreting the biological data they obtain from these experiments and observations. The laboratories are an integral part of the courses; to receive a grade in a Common Core course, students must complete the laboratory exercises associated with that course even if the course is taken as an elective. NOTE: Some courses include laboratories that require the use of animals; students who have concerns about using animals in laboratories are urged to discuss these concerns with the instructors before registering for those courses.

Placement.
Students who score 4 or 5 on the AP test in biology, or who perform well on the College's optional placement test in the biological sciences, will be excused from the Common Core biological sciences requirement. Students who place out of the biology Common Core and who subsequently concentrate in the biological sciences are required either to complete one of the Biological Sciences 170s, 180s, or 190s sequences as part of their concentration requirement or demonstrate their knowledge by special examination.

Accreditation.
Students who place out of the Common Core biology sequence may elect to take an accreditation examination in one or more of the five areas included in the Biological Sciences 170s, 180s, and 190s sequences. Credit is granted upon satisfactory completion of an examination designated by the BSCD master. No laboratory requirements can be met by accreditation examinations except by special petition with accompanying documentation. Accreditation for 200-level biology courses may also be granted to students at any time after completion of a 100-level Common Core biology sequence and upon satisfactory completion of the same examination. No examination for accreditation may be taken more than once.

Students are urged to consult the BSCD senior adviser immediately if they feel that they can qualify for accreditation.

The Biological Sciences Concentration Program

The goals of the biological sciences concentration are to give students an understanding of currently accepted concepts in biology and the experimental support for these concepts, and an appreciation of the gaps in our current understanding and the opportunities for new research in this field. The concentration is designed to prepare students for graduate or professional study in the biological sciences and for careers in biology. As in the Common Core, emphasis is placed on introducing students to the diversity of subject matter and methods of investigation in the biological sciences.

The requirements for a B.A. in the biological sciences include the following:

1. The Common Core. To prepare themselves for more advanced work in biology, prospective biological sciences concentrators should take Chemistry 111-112-113 or higher to satisfy the Common Core requirement in the physical sciences and Mathematics 131-132 or higher to fulfill the mathematics requirement. Students who concentrate in the biological sciences may satisfy their Common Core requirement in the biological sciences with one of the Biological Sciences 170s, 180s, or 190s sequences or with any other Common Core biological sciences sequence. The five courses from one of the 170s, 180s, or 190s sequences, as well as a course in biochemistry, constitute a six-course requirement designed to prepare students for advanced study in the concentration. Students who complete a Common Core sequence other than one of the 170s, 180s, or 190s sequences should consult with the BSCD master and senior adviser about the courses to take to meet the concentration requirements.

2. Other Science Courses. Biological sciences concentrators must complete two quarters of organic chemistry (Chemistry 217-218 or 220-221), two quarters of physics (Physics 121-122 or higher), one additional quarter of mathematics (Mathematics 133 or higher) or statistics (Statistics 220), and one additional course in mathematics or physical sciences.

3. Advanced-Level Biology Courses.

a. Biological Sciences 170s, 180s, or 190s sequences. Students who take three of these courses to fulfill the Common Core biological sciences requirement will take the remaining two courses as part of their concentration program.

b. Biological Sciences 200 (Introduction to Biochemistry).

c. Students may fulfill the concentration requirement in the biological sciences with a general program in the biological sciences or with a specialization program. In either case, five additional courses in the biological sciences or related disciplines at the 200-level or above are required. Students who choose the general program in biological sciences will take one course in either cell, molecular, genetics, or developmental biology (one of Biological Sciences 207-229), one in organismal biology (one of Biological Sciences 232-243 or 260), and one in ecology, evolutionary biology, population genetics, or biological diversity (one of Biological Sciences 248-256). Students who choose to specialize will take three courses designated by their specialization program. All students will have two electives within the concentration. Biological Sciences 199, 297, 298, and 299 may not be used to satisfy this requirement.

4. Laboratories. In addition to the laboratory required for Biological Sciences 200, two other 200-level biological science courses must have a laboratory. Courses used to satisfy the laboratory requirement need not be the same courses used to satisfy other concentration requirements, but they can be. The two laboratory courses must be in different areas of the biological sciences.

Summary of Requirements

General Chem 111-112-113 or 121-122-123

Education
Math 131-132, 151-152, or 161-162

(a) BioSci 170s, 180s, or 190s sequence (three courses) or
(b) any other biological sciences Common Core sequence*

Concentration For students who choose option (a) above:

2 Completion of BioSci 170s, 180s, or 190s sequences
1 biochemistry (BioSci 200)
5 courses in the biological sciences or related disciplines at the 200-level or above
2 organic chemistry (Chem 217-218 or 220-221)
2 Phys 121-122, 131-132, or 141-142
1 Math 133, 153, or 163, or Stat 220
1 additional course in mathematics or

physical sciences

14

*Students who choose option (b) in the summary above should consult with the BSCD master and senior adviser about the courses to take to meet the concentration requirements. Generally they should expect to complete all or part of one of the five-course 170s, 180s, or 190s sequences as part of their concentration courses.

Specialization Programs in the Biological Sciences

Specialization in Cellular and Molecular Biology.
Students who complete the following specific courses and fulfill the following lab requirements en route to completing the biological sciences concentration requirements will be recognized as having completed a specialization in the area of cellular and molecular biology.

The following requirements must be met:

Courses
1. organic chemistry (Chem 219 or 222). (Fulfills the concentration requirement for one additional course in mathematics or physical sciences.)

2. The following upper level courses in biological sciences:

a. one course in the area of cell biology (BioSci 207 or 209)

b. one course in the area of genetics (BioSci 217 or 239)

c. one course in the area of developmental biology (BioSci 226/7 or 229)

Laboratory Two of these courses must be laboratory courses unless either BioSci 215 or 216 is taken as an elective.

The specialization in cellular and molecular biology is administered by the Department of Molecular Genetics and Cell Biology.

Specialization in Ecology and Evolution.
Students who complete the following course work and fulfill the requirements of the senior honors paper en route to completing the biological sciences concentration requirement will be recognized as having completed a specialization in ecology and evolution. Each student will be assigned a faculty adviser based upon the student's particular topic of interest. The faculty adviser will recommend specific courses necessary to meet the requirements (see following section) and facilitate placing the student in a research laboratory to design and complete the original research necessary for the senior honors project. The specialization is recommended for those students interested in pursuing graduate work in the field or laboratory sciences of ecology, evolution, population genetics, and behavior.

The following requirements must be met:

Courses
1. three quarters of calculus and two quarters of statistics (in lieu of physics requirement),

2. three upper level courses in the biological sciences as recommended by the faculty adviser and departmental chair from a menu of courses in ecology, evolution, genetics, and behavior

Laboratory
Completion of original research in the lab and under the guidance of a member of the ecology and evolution faculty, qualifying the student for a senior honors paper.

The specialization in ecology and evolution is administered by the Department of Ecology and Evolution. Please contact Dr. Michael J. Wade (chairman) at 702-9638 or mwade@pondside.uchicago.edu.

Specialization in Neuroscience.
Students who complete the following requirements en route to completing the biological sciences concentration requirements will be recognized as having completed a specialization in neuroscience. Students who elect to specialize will be assigned to a faculty adviser (Dorothy Hanck, Daniel Margoliash, or Martha McClintock), who will be available to advise them on classes that would best accomplish successful completion of the specialization and help them identify laboratories in which they can carry out their individual research projects.

Required courses include

BioSci 210 (Developmental Biopsychology)
BioSci 211 (Systems and Behavioral Neurobiology)
BioSci 212 (Cellular Neurobiology)

In addition, in consultation with their faculty adviser, each student will choose two electives from the following list:

BioSci 214 (Developmental Neurobiology)
BioSci 230 (Ion Channels)
BioSci 231 (Nonlinear Dynamics for Neuroscience and Biopsychology)
BioSci 258 (Neuropharmacology)
BioSci 268 (Neuropsychopharmacology)
BioSci 270 (Conquest of Pain)
Biopsy 215 (Brain Asymmetry)
Biopsy 231 (Developmental Neuropsychology)
Biopsy 280 (Sensation and Perception)
Biopsy 287 (Connectionist Modeling I)
Biopsy 291 (Connectionist Modeling II)

Students are strongly encouraged to carry out individual guided research, to participate in the honors research program, and to attend neurobiology/biopsychology related research seminars. The specialization in neuroscience is administered by the Committee on Neurobiology.

Research Courses and Departmental Honors.
Students are strongly encouraged to carry out individual guided research in an area of their interest. Students who choose to carry out research can receive academic credit for their work and have the opportunity to graduate with special honors. A student may propose an arrangement with any faculty member in the Biological Sciences Collegiate Division to sponsor and supervise his or her research on an individual tutorial basis; the student's activities are arranged by mutual agreement between the student and the sponsor. Students can learn about research opportunities and potential faculty sponsors from the College Research Opportunities Program (CROP) Directory, which is available on the World Wide Web (http://www-college.uchicago.edu/crop/crop.html).

Students can receive academic credit for their research by enrolling in research courses. Two such courses are available: Biological Sciences 199 (Introduction to Research) and Biological Sciences 299 (Advanced Research in the Biological Sciences). Biological Sciences 199 is intended for students who wish to have a research experience but are not conducting an approved honors research project; it may be elected for up to three quarters. Biological Sciences 299 is for advanced students, including but not limited to those who wish to be candidates for special honors. Registration for either Biological Sciences 199 or Biological Sciences 299 requires permission of the student's research sponsor and the chairman of the Undergraduate Research and Honors Committee. To register for Biological Sciences 299, students must submit a written research proposal to the chairman of the Undergraduate Research and Honors Committee. A registration form for these courses may be obtained from your adviser. Students must complete this form before registering for research courses. Both Biological Sciences 199 and Biological Sciences 299 are graded P/F; neither course can be used to fulfill concentration requirements in the biological sciences.

Students who take Biological Sciences 299 are required to participate in a biweekly senior honors forum beginning in autumn quarter. In addition, students who wish to receive special honors must participate in Biological Sciences 298 (Undergraduate Research Seminar) in their senior year. Students who are unable to take Biological Sciences 298 for credit may audit this course with the approval of the chairman of the Undergraduate Research and Honors Committee. In the honors forum and seminar, students report on and discuss their research. Students who take Biological Sciences 199 are encouraged to attend the forum and seminar but will normally not present their work. Biological Sciences 298 is offered in the spring quarter; students who cannot take this course because they plan to graduate in a different quarter should contact the chairman of the Undergraduate Research and Honors Committee at least three months before their expected graduation to arrange for an equivalent experience. Biological Sciences 298 is also graded P/F and may not be used to fulfill concentration requirements in the biological sciences.

Grading.
Students must receive quality (letter) grades in all fourteen courses in the concentration.

Combined Bachelor of Arts/Master of Science Degree Program in Biochemistry and Molecular Biology

This program is designed for those students who, early in their academic careers, decide to pursue graduate study in biology at the molecular level. It differs from the usual concentration in the biological sciences in that it requires a central core of graduate courses in biochemistry and molecular biology. These in turn require a background in physical chemistry.

This program requires the completion of forty-eight credits of course work rather than the forty-two required for the B.A. (NOTE: Fifty-one credits--forty-two for the B.A. and nine for the M.S.--are required; however, up to three upper-level course credits can be double-counted for both degrees.) Thus, the program is difficult in terms of the course load and also requires payment of graduate-level tuition during the senior year. In general, the program is suitable only for students who enter with placement or advanced placement credit or who complete some of their work during the summer.

A departmental oral examination on a thesis describing the candidate's original research contribution is required for the degree. Students are expected to (1) spend elective time and/or a summer residency in a laboratory working on a research problem in biochemistry or molecular biology, and (2) write a formal thesis describing their research and reviewing the field. The oral examination based on that thesis must be taken during the last quarter of their registration.

Students will not be admitted formally to the program until their third year. Applications are made through the Department of Biochemistry and Molecular Biology. Normally only students with grade point averages above 3.0 qualify, although exceptions may be made under special circumstances.

Questions about the program may be addressed to the adviser in the Department of Biochemistry and Molecular Biology (Marvin Makinen, CLSC 359, 702-1080).

Faculty

JEANNE ALTMANN, Professor, Department of Ecology & Evolution, Committees on Biopsychology and Evolutionary Biology, and the College; Chairman, Committee on Evolutionary Biology

STEVAN J. ARNOLD, Professor, Department of Ecology & Evolution, Committees on Evolutionary Biology and Genetics, and the College

R. DEAN ASTUMIAN, Assistant Professor, Departments of Biochemistry & Molecular Biology and Surgery and the College

JUDITH AUSTIN, Assistant Professor, Department of Molecular Genetics & Cell Biology, Committee on Developmental Biology, and the College

JOY M. BERGELSON, Assistant Professor, Department of Ecology & Evolution and the College

RUDIGER BIELER, Lecturer, Committee on Evolutionary Biology and the College

ANDREW BIEWENER, Associate Professor, Department of Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the College; Chairman, Department of Organismal Biology & Anatomy

JEFFREY A. BLUESTONE, Professor, Ben May Institute, Department of Pathology, Committee on Immunology, and the College; Chairman, Committee on Immunology

JAMES E. BOWMAN, M.D., Professor Emeritus, Departments of Pathology and Medicine, Committees on African & African-American Studies and Genetics, and the College

DONN BRANSTRATOR, Lecturer in the College

MALCOLM J. CASADABAN, Associate Professor, Departments of Biochemistry & Molecular Biology and Molecular Genetics & Cell Biology, Committees on Genetics and Virology, and the College

BRIAN CHARLESWORTH, George Wells Beadle Distinguished Service Professor, Department of Ecology & Evolution, Committees on Evolutionary Biology and Genetics, and the College

DEBORAH CHARLESWORTH, Professor, Department of Ecology & Evolution, Committees on Evolutionary Biology and Genetics, and the College

BARRY CHERNOFF, Lecturer, Committee on Evolutionary Biology and the College

KWEN-SHENG CHIANG, Associate Professor, Department of Molecular Genetics & Cell Biology, Committee on Genetics, and the College

LIANE COCHRAN-STAFIRA, Lecturer in the College

THOMAS COLTON, Senior Lecturer in the College

JERRY COYNE, Professor, Department of Ecology & Evolution, Committees on Evolutionary Biology and Genetics, and the College

MARY CRANE, Lecturer in the College

RONALD DUBREUIL, Assistant Professor, Department of Pharmacological & Physiological Sciences, Committee on Cell Physiology, and the College

WOLFGANG EPSTEIN, M.D., Professor, Departments of Biochemistry & Molecular Biology and Molecular Genetics & Cell Biology, Committee on Genetics, and the College

ROCHELLE EASTON ESPOSITO, Professor, Department of Molecular Genetics & Cell Biology, and the College; Chairman, Committee on Genetics

MARTIN E. FEDER, Professor, Department of Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the College

EDWIN FERGUSON, Assistant Professor, Department of Molecular Genetics & Cell Biology, Committees on Developmental Biology and Genetics, and the College

FRANK W. FITCH, M.D., Albert D. Lasker Professor, Department of Pathology, Committees on Developmental Biology and Immunology, and the College; Director, Ben May Institute

JOHN FLYNN, Lecturer, Committee on Evolutionary Biology and the College

MICHAEL FONSTEIN, Research Associate (Assistant Professor), Department of Molecular Genetics & Cell Biology; Lecturer in the College

LINCOLN E. FORD, M.D., Associate Professor, Department of Medicine and the College

HARRY A. FOZZARD, M.D., Otho S. A. Sprague Distinguished Service Professor, Departments of Medicine and Pharmacological & Physiological Sciences, Committee on Cell Physiology, and the College; Chairman, Department of Pharmacological & Physiological Sciences

HERBERT C. FRIEDMANN, Associate Professor, Department of Biochemistry & Molecular Biology and the College

EDWARD D. GARBER, Professor Emeritus, Department of Ecology & Evolution, Committee on Genetics, and the College

LAWRENCE M. GARTNER, M.D., Professor, Department of Pediatrics and the College

GODFREY S. GETZ, M.D., Donald N. Pritzker Professor, Departments of Biochemistry & Molecular Biology, Medicine, and Pathology, and the College; Chairman, Department of Pathology

ANN D. GOLDBLATT, Lecturer, Department of Medicine and the College

LARRY GOLDMAN, M.D., Associate Professor, Department of Psychiatry and the College

EUGENE GOLDWASSER, Alice Hogge and Arthur A. Baer Professor Emeritus, Department of Biochemistry & Molecular Biology, Committee on Developmental Biology, and the College; Chairman, Department of Biochemistry & Molecular Biology

DANIEL GOTTSCHLING, Associate Professor, Department of Molecular Genetics & Cell Biology, Committee on Genetics, and the College

GEOFFREY GREENE, Professor, Ben May Institute, Department of Biochemistry & Molecular Biology and the College

RICHARD GUNDERMAN, Lecturer in the College

DOROTHY A. HANCK, Assistant Professor, Department of Medicine, Committees on Cell Physiology and Neurobiology, and the College

ROBERT HASELKORN, Fanny L. Pritzker Distinguished Service Professor, Departments of Biochemistry & Molecular Biology, Chemistry, and Molecular Genetics & Cell Biology, Committees on Developmental Biology, Genetics, and Virology, and the College

ALFRED HELLER, M.D., Professor, Department of Pharmacological & Physiological Sciences and the College

PHILIP C. HOFFMANN, Professor, Department of Pharmacological & Physiological Sciences and the College

JAMES A. HOPSON, Professor, Department of Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the College

LYNNE HOUCK, Research Associate (Associate Professor), Department of Ecology & Evolution; Lecturer in the College

JOSEPH JARABAK, M.D., Associate Professor, Department of Medicine and the College

ROBERT JOSEPHS, Professor, Department of Molecular Genetics & Cell Biology and the College

BRIAN KEITH, Assistant Professor, Department of Molecular Genetics & Cell Biology, Committees on Developmental Biology and Genetics, and the College

ARTHUR F. KOHRMAN, M.D., Professor, Department of Pediatrics and the College

CLAIRE KOHRMAN, Research Associate (Associate Professor), Center for Health Administration Studies and Department of Pediatrics; Lecturer in the College

MARTIN KREITMAN, Associate Professor, Department of Ecology & Evolution, Committees on Evolutionary Biology and Genetics, and the College

JOHN KRUPER, Director, Academic Computing, Division of the Biological Sciences; Lecturer in the College

ROBERT KUSHNER, M.D., Assistant Professor, Department of Medicine and the College

MICHAEL C. LABARBERA, Professor, Departments of the Geophysical Sciences and Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the College

GAYLE LAMPPA, Associate Professor, Department of Molecular Genetics & Cell Biology, Committee on Genetics, and the College

JOHN LANTOS, M.D., Associate Professor, Departments of Medicine and Pediatrics and the College

MATHEW LEIBOLD, Assistant Professor, Department of Ecology & Evolution, Committee on Evolutionary Biology, and the College

SUSAN LINDQUIST, Professor, Howard Hughes Medical Institute, Department of Molecular Genetics & Cell Biology, Committees on Developmental Biology and Genetics, and the College

PHILIP E. LLOYD, Associate Professor, Department of Pharmacological & Physiological Sciences, Committees on Cell Physiology and Neurobiology, and the College; Chairman, Committee on Neurobiology

R. ERIC LOMBARD, Associate Professor, Department of Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the College

ANTHONY P. MAHOWALD, Louis Block Professor, Department of Molecular Genetics & Cell Biology, Committees on Developmental Biology, Genetics, and Neurobiology, and the College; Chairman, Department of Molecular Genetics & Cell Biology and Committee on Developmental Biology

MARY MAHOWALD, Professor, Department of Obstetrics & Gynecology and the College

MARVIN W. MAKINEN, Professor, Department of Biochemistry & Molecular Biology and the College

DANIEL MARGOLIASH, Associate Professor, Departments of Organismal Biology & Anatomy and Psychology, Committees on Biopsychology and Neurobiology, and the College

ALVIN MARKOVITZ, Professor, Department of Biochemistry & Molecular Biology, Committee on Genetics, and the College

TERENCE E. MARTIN, Professor, Department of Molecular Genetics & Cell Biology, Committees on Genetics and Immunology, and the College

MARK MARTINDALE, Assistant Professor, Department of Organismal Biology & Anatomy, Committees on Developmental Biology and Neurobiology, and the College

MANISHA H. MASKAY, Assistant Professor, Department of Medicine and the College

JANE MASTERSON, Lecturer in the College

MARTHA K. MCCLINTOCK, Professor, Department of Psychology, Committees on Evolutionary Biology and Neurobiology, and the College; Chairman, Committee on Biopsychology

RIMA MCLEOD, M.D., Lecturer, Department of Medicine, Committee on Immunology, and the College

STEPHEN C. MEREDITH, M.D., Associate Professor, Department of Pathology and the College

LAURENS METS, Associate Professor, Department of Molecular Genetics & Cell Biology, Committee on Genetics, and the College

JAMES MILLER, Associate Professor, Departments of Molecular Genetics & Cell Biology and Pathology, Committees on Developmental Biology and Immunology, and the College

MALKA MOSCONA, Associate Professor in the College

JONATHAN MOSS, M.D., Professor, Department of Anesthesiology & Critical Care and the College

GREGORY H. MUELLER, Lecturer, Committee on Evolutionary Biology and the College

MARK MUSCH, Research Associate (Assistant Professor), Department of Medicine; Lecturer in the College

JAMES NACHMAN, M.D., Associate Professor, Department of Pediatrics and the College

THOMAS NAGYLAKI, Professor, Department of Ecology & Evolution, Committees on Evolutionary Biology and Genetics, and the College

MATTHEW NITECKI, Curator, Committee on Evolutionary Biology; Lecturer in the College

AVERTANO NORONHA, M.D., Associate Professor, Department of Neurology, Committee on Immunology, and the College

CAROLE OBER, Associate Professor, Department of Obstetrics & Gynecology, Committees on Evolutionary Biology and Genetics, and the College

SUJATA PATEL, Lecturer in the College

BRUCE PATTERSON, Lecturer, Committee on Evolutionary Biology and the College

ROBERT L. PERLMAN, M.D., Professor, Departments of Pediatrics and Pharmacological & Physiological Sciences, Committees on Cell Physiology and Neurobiology, and the College

CATHY A. PFISTER, Assistant Professor, Department of Ecology & Evolution and the College

ROSAMOND V. POTTER, Senior Lecturer in the College

STEPHEN PRUETT-JONES, Associate Professor, Department of Ecology & Evolution, Committee on Evolutionary Biology, and the College

JOSé QUINTANS, M.D., Professor, Department of Pathology and the College; Master, Biological Sciences Collegiate Division; Associate Dean, Division of Biological Sciences and the College

HYMAN ROCHMAN, M.D., Associate Professor, Department of Pathology and the College

BERNARD ROIZMAN, Joseph Regenstein Distinguished Service Professor, Departments of Biochemistry & Molecular Biology and Molecular Genetics & Cell Biology, Committees on Genetics and Virology, and the College; Chairman, Committee on Virology

MARSHA R. ROSNER, Professor, Ben May Institute, Department of Pharmacological & Physiological Sciences, Committee on Cell Physiology, and the College; Chairman, Committee on Cancer Biology

LUCIA ROTHMAN-DENES, Professor, Department of Molecular Genetics & Cell Biology, Committee on Genetics, and the College

MANFRED D. E. RUDDAT, Associate Professor, Department of Ecology & Evolution, Committee on Developmental Biology, and the College; Associate Dean of the College; Senior Adviser, Biological Sciences Collegiate Division

DALE A. SCHOELLER, Associate Professor, Department of Medicine and the College; Chairman, Committee on Human Nutrition & Nutritional Biology

SIDNEY SCHULMAN, M.D., Ellen C. Manning Professor Emeritus, Division of the Biological Sciences and the College

GEBHARD F. B. SCHUMACHER, M.D., Professor Emeritus, Department of Obstetrics & Gynecology and the College

PAUL T. SCHUMACKER, Professor, Department of Medicine and the College

LEWIS S. SEIDEN, Professor, Departments of Pharmacological & Physiological Sciences and Psychiatry, Committee on Neurobiology, and the College

J. JOHN SEPKOSKI, JR., Professor, Departments of the Geophysical Sciences and Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the College

PAUL SERENO, Associate Professor, Department of Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the College

JAMES SHAPIRO, Professor, Department of Biochemistry & Molecular Biology, Committee on Genetics, and the College

PETRA SIERWALD, Lecturer in the College

ELLEN SIMMS, Assistant Professor, Department of Ecology & Evolution, Committee on Evolutionary Biology, and the College

PATRICIA A. SOLTYS, Lecturer in the College

THEODORE L. STECK, M.D., Professor, Departments of Biochemistry & Molecular Biology and Molecular Genetics & Cell Biology, Committee on Cell Physiology, and the College

GLENN D. STEELE, M.D., Dean, Division of the Biological Sciences and the Pritzker School of Medicine and Vice President for Medical Affairs

DONALD F. STEINER, M.D., A. N. Pritzker Professor, Howard Hughes Medical Institute, Departments of Biochemistry & Molecular Biology and Medicine, and the College

URSULA STORB, Professor, Departments of Molecular Genetics & Cell Biology and Pathology, Committees on Genetics, Developmental Biology, and Immunology, and the College

FRANCIS H. STRAUS II, M.D., Professor, Department of Pathology and the College

LORNA P. STRAUS, Professor, Department of Organismal Biology & Anatomy and the College

BERNARD S. STRAUSS, Professor, Department of Molecular Genetics & Cell Biology, Committee on Genetics, and the College

PAUL STRIELEMAN, Lecturer in the College

DON R. SWANSON, Professor, Division of the Humanities and the College

KEVIN SWIER, Lecturer in the College

SARA SZUCHET, Professor, Department of Neurology, Committee on Neurobiology, and the College

EDWIN W. TAYLOR, Louis Block Professor, Departments of Biochemistry & Molecular Biology and Molecular Genetics & Cell Biology, Committee on Cell Physiology, and the College

FRANK K. THORP, Associate Professor, Department of Pediatrics, Committee on Human Nutrition & Nutritional Biology, and the College

RUSSELL H. TUTTLE, Professor, Department of Anthropology, Committee on Evolutionary Biology, Morris Fishbein Center for the History of Science & Medicine, and the College

PHILIP S. ULINSKI, Professor, Department of Organismal Biology & Anatomy, Committee on Neurobiology, and the College

LEIGH M. VAN VALEN, Professor, Department of Ecology & Evolution, Committees on Conceptual Foundations of Science, Evolutionary Biology, and Genetics, and the College

MARION VERP, Associate Professor, Department of Obstetrics & Gynecology and the College

MITCHEL VILLEREAL, Professor, Department of Pharmacological & Physiological Sciences, Committee on Cell Physiology, and the College; Chairman, Committee on Cell Physiology

MICHAEL J. WADE, Professor, Department of Ecology & Evolution, Committees on Evolutionary Biology and Genetics, and the College; Chairman, Department of Ecology & Evolution

NORMA E. WAGONER, Professor, Department of Organismal Biology & Anatomy and the College; Dean of Students, Division of the Biological Sciences and the Pritzker School of Medicine

MARK WESTNEAT, Lecturer, Department of Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the College

HUGH R. WILSON, Professor, Department of Ophthalmology & Visual Science, Committee on Neurobiology, and the College

WILLIAM WIMSATT, Professor, Department of Philosophy, Committees on Conceptual Foundations of Science, Evolutionary Biology, and General Studies in the Humanities, Morris Fishbein Center for the History of Science & Medicine, and the College

J. TIMOTHY WOOTTON, Assistant Professor, Department of Ecology & Evolution, Committee on Evolutionary Biology, and the College

RADOVAN ZAK, M.D., Professor, Departments of Medicine, Organismal Biology & Anatomy and Pharmacological & Physiological Sciences, Committee on Cell Physiology, and the College

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